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Compound

Pectin methylesterase

The enzyme that strips methyl groups from pectin, exposing the charged sites that calcium bridges — which is the reaction the whole of keeving is built on.

Also called PME, Pectinesterase.

Class
Enzymes
Formula
Not a single molecule — see below
How often it matters
Central to how cider behaves

What it does in cider

What forms it

Processes that put this compound into the drink, or increase how much of it is there.

What removes or limits it

Processes that reduce it, hold it below a threshold, or stop it forming in the first place.

Faults it is implicated in

Being implicated is not the same as being a fault. Several of the compounds on this site are ordinary constituents of a sound cider and define a named fault only above a concentration.

About Pectin methylesterase

Pectin methylesterase does one thing: it hydrolyses the methyl ester groups that cap many of the galacturonic acid units in a pectin chain. The methanol released is the trivial part of the reaction. The important part is what is left behind — a free carboxyl group, negatively charged at cider pH, at each position the enzyme has visited.

A pectin chain with enough of these free sites can be cross-linked. Calcium ions, each carrying two positive charges, bridge carboxyl groups on adjacent chains, and once enough bridges form the dissolved polymer becomes a coherent three-dimensional gel. This is the défécation of French practice and the keeve of English practice, and the whole apparatus of it — cold juice, calcium salt, slow incipient fermentation, days of patience — exists to give this one enzyme time to do its work at a temperature that stops yeast doing anything faster.

The difficulty in practice is that native activity varies enormously with cultivar, with maturity and with how the fruit was stored, so a traditional keeve either works or does not, and generations of makers have described it as unpredictable. Commercial keeving preparations supply the enzyme directly and make the outcome repeatable, at the cost of the thing being wholly the fruit’s doing.

The same enzyme is a nuisance in other contexts. In cloudy juice production it de-esterifies pectin gradually in the pack, and the calcium naturally present then forms a gel or a haze in a product that was stable at bottling — which is why juice intended to stay cloudy is usually pasteurised warm enough to kill the enzyme as well as the microorganisms.

Related compounds

Compounds it is formed from, converted into, confused with, or routinely met alongside.

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Questions readers ask about the things this page mentions. Each one goes to the section that answers it rather than to a page written to receive the question.

Where to go next

Sources

What this page rests on. Where a source is marked as registered rather than read, CiderHQ is recording that the body is authoritative on the subject without claiming to have worked through the document itself. See our evidence policy for what each state means.